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Ellie and Manny are expecting their first child, and Manny is obsessed with making life perfect and safe for the family. He is shown bringing a shell full of water to Ellie believing that she is giving birth, although this is just a false alarm. At the same time, Diego finds himself unable to catch an antelope he has been stalking. The antelope starts to taunt him. Believing that he is losing his predatory Sid with his kids
Sid with dinosour eggs as a tiger, he decides to leave the herd after a talk with Manny. Sid grows jealous of Manny and Ellie and �adopts� three apparently abandoned eggs that he finds in an icy underground cavern. Manny tells him to put them back, but Sid instead looks after the eggs, which hatch into baby Tyrannosaurus the next morning.
Although Sid tries his best to raise the three dinosaurs, their rambunctious behavior scares away all the other animals� young and ruins a playground Manny built for Ellie�s baby. The mother dinosaur soon returns and carries both Sid and her youth underground, with Diego in pursuit. Manny, Ellie, Crash, and Eddie follow as well and discover that the icy cavern leads to a vast jungle populated by dinosaurs thought to be extinct. Here, an Ankylosaurus threatens the herd despite Diego�s efforts to fend it off; they are saved from a further crowd of angry reptiles (including Kentrosaurus, Brachiosaurus, Maiasaura, and Pachycephalosaurus) by a one-eyed, slightly crazed weasel named Buckminster, or Buck.
Buck appears
Buck appears to help the herd.
Buck has been living in this jungle for some time and chasing Rudy (a giant white Baryonyx that is this world�s biggest dinosaur), intending to avenge the eye he lost to it. He agrees to lead the herd through the jungle�s perils to Lava Falls, where �Momma� has taken Sid and her babies. Soon Manny and Diego are captured by a carnivorous plant, but thanks to Buck, they are freed. In the meantime, Sid and the mother try to outdo each other in feeding the offspring; he loses this contest, but is soon welcomed into the family regardless.
That night, while the herd was resting Manny had a bad dream of Ellie all alone in a jungle and being killed by Rudy. After he wakes up, Ellie was still sleeping; however, she woke up, too, and Manny explains to her about his nightmare. Because Ellie was in the nightmare, she explains the most dangerous place in the world is bigger than her and Manny. Buck hears that if Manny was nicer to Sid, they wouldn't be searching for him. Buck says that Manny risked his own life and his unborn baby. That makes Manny a friend and a hero.
The next day, however, Rudy attacks the (dinosaur) family and Sid is knocked onto a loose rock slab that is floating on a river of lava and about to plummet over the falls.
As the herd moves toward Lava Falls, Ellie goes into labor and a Guanlong pack strikes, causing a rock slide that separates her from Manny and Diego. Manny doubles back to protect her and Diego fends off further attacks, while Buck takes Crash and Eddie ahead to rescue Sid. Just as the sloth goes over the falls, the trio swoops in on a commandeered Pteranodon and saves his life (after being chased by some Quetzalcoatlus). Manny reaches Ellie just as their baby is born, a girl they decide to name Peaches (the codeword they had chosen for Ellie to use if she went into labor during the trip). Sid is saddened at the fact that he never had a chance to say goodbye to "his" children as he is reunited with the herd.
Before they can exit the jungle, Rudy attacks at full force; Buck helps the others ensnare him briefly, but he escapes and resumes his onslaught. The herd is saved by the timely arrival of Momma, who charges at Rudy and knocks him off a cliff. As she and her children wish Sid well, Buck � now without a purpose in life since Rudy is gone � decides to join the herd and live on the surface. However, a distant roar tells him that Rudy is still alive; he changes his mind and sends the herd home, he destroys the path to the underground jungle at the same time so that no one can go down there. Manny and Ellie welcome Peaches into their frozen world and admit that Sid did a good job looking after Momma's children. Diego decides to remain with the herd, while Buck stays where he wants to be: underground, matching wits with Rudy.?
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